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The film begins in an abandoned river boat in Devil's Bayou, where orphan Penny drops a message in a bottle containing a plea for help into the river. The bottle is carried out to sea and washes up in New York, where it is recovered by the Rescue Aid Society. The Hungarian representative, Miss Bianca, volunteers to accept the case and chooses the janitor Bernard as her co-agent. The two visit Morningside Orphanage, where Penny lived, and meet an old cat named Rufus. He tells them about a wicked woman named Madame Medusa who once tried to lure Penny into her car and may have abducted Penny this time.The mice travel to Medusa's pawn shop, where they discover that she and her partner Mr. Snoops are on a quest to find the world's largest diamond, the Devil's Eye. With the help of an albatross named Orville, and a dragonfly named Evinrude, the mice follow Medusa and Mr. Snoops to the bayou. There, they learn that Penny was captured to enter a hole that leads down into the pirates' cave where the Devil's Eye is located.
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